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> Social order works better than the alternatives, in which parasites win.

You're part of the way there. Individual units may play "Cooperate" or "Defect". But iterated prisoner's dilemma is static, and this is dynamic. So "Defect" can end up being advantageous for every individual and after that, it's integrated into the ruleset. You're entering the office, you bump into a colleague in the parking lot, you swipe your badge and the door opens, he comes in with you without swiping, you hold the door. Either you've cooperated or you've defected.