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in the mammalian articulatory system, it was the development of costal breathing that allowed the laryngeal muscles to become free as an articulatory organ. this also coincided with rearranging the leverage and musculature in the jaw which allowed the repurposing of the bones in the jaw to form the bones of the middle ear. It is believed that these adaptations could have been subject to joint selection for hearing, chewing, and vocalization. So the slugs developing the ability to scream alongside the ability to eat meat tracks, at least loosely, the course of mammalian evolution for some related body systems #monsterdon